Issue: Colour in Harmony

I've encountered an issue with Toon Boom Harmony. The colour I've applied to my film (as visible in the screen recording below) doesn't show up when I export or render the video clip. Any areas that contain colour show up as black. 

Screen Recording of Scene 1 Shot 1 - Coloured

This is what the colour should look like, as it appears in Harmony. 

Export Showing Colour Issue

When I exported the clip above, this is what it appeared as. 

Export with Lineart and Coloured Hands

I removed the colour (except on the hands, which I forgot), and kept only the lineart. We can see that the issue is not in the rest of the film, and only the colour is showing up as black. 

I've been colouring with harmony's vector drawing. One of my peers is drawing in the same way and her colour shows up, so I'm not sure what I'm doing wrong.